Constance Marten, 35, and Mark Gordon, 48, will make their first court appearance in Crawley, West Sussex, today.
The couple have been charged with gross negligence manslaughter after the remains of a baby were found in Brighton.
Constance Marten, 35, and Mark Gordon, 48, who had been missing since 5 January, were arrested in the city on Monday after being spotted by a member of the public in a shop.
They were found without their newborn child.
A major search involving more than 200 police officers was carried out for the infant in allotments and woodland in and around Brighton.
Police said on Wednesday that the remains of a baby had been found in a wooded part of the Hollingbury area.
Marten, who comes from an aristocratic family, and her boyfriend were charged with gross negligence manslaughter on Thursday evening, said the Metropolitan Police.
They were also charged with concealing the birth of a child and perverting the course of justice.
The pair will appear at Crawley Magistrates' Court today.
Police had been looking for Marten and Gordon for nearly two months, since their car caught fire on the motorway near Bolton.
There were sightings of them in various locations around the UK - most recently in East Sussex.
Earlier on Thursday, police said they did not yet know the gender of the baby and that it had been dead for several weeks.
The baby has not yet been formally identified and a post-mortem is due to be carried out today to discover how it died.
People in Brighton have been leaving flowers, messages and cuddly toys on a road close to where the child was found.
The Bishop of Lewes also led a candlelight vigil at a nearby church on Thursday night.
(c) Sky News 2023: Constance Marten and Mark Gordon charged with gross negligence manslaughter
More 'Live' Bus Information Screens To Be Installed In West Sussex
Rare Bee Found In Lewes
Event Bookings Go Live For Worthing Festival ‘24
Man Charged After Throwing Brick Through Brighton Restaurant Window
Peacehaven Man Disqualified Over High-Speed A27 Pursuit Near Brighton
Appeal After PCSO Assaulted In Uckfield
Man Charged With Rape Of Teenage Girl In Newhaven
Overnight Closures For A22 Forest Row Road Improvements
Brighton Defence Manufacturer's Controversial Planning Application Likely To Be Heard
Over £2.4m Approved For Worthing School Support Centre